1. Zhang, Hongdian, Han, Youming, Xiao, Wanyi, Tang, Peng, Yu, Zhentao. 2023. USP4 promotes the proliferation, migration, and invasion of esophageal squamous cell carcinoma by targeting TAK1. In Cell death & disease, 14, 730. doi:10.1038/s41419-023-06259-0. https://pubmed.ncbi.nlm.nih.gov/37949874/
2. Zhou, Jiangqiao, Qiu, Tao, Wang, Tianyu, Zhang, Long, Zou, Jilin. 2019. USP4 deficiency exacerbates hepatic ischaemia/reperfusion injury via TAK1 signalling. In Clinical science (London, England : 1979), 133, 335-349. doi:10.1042/CS20180959. https://pubmed.ncbi.nlm.nih.gov/30622220/
3. Hu, Chuanxiang, Zhang, Wei, Jia, Yongsheng, Hao, Chengfei, Yu, Yang. 2024. USP4 promotes PTC progression by stabilizing LDHA and activating the MAPK and AKT signaling pathway. In Aging, 16, 12850-12865. doi:10.18632/aging.206108. https://pubmed.ncbi.nlm.nih.gov/39393052/
4. Zhong, Ming, Jiang, Qi, Jin, Ronghui. 2018. USP4 expression independently predicts favorable survival in lung adenocarcinoma. In IUBMB life, 70, 670-677. doi:10.1002/iub.1755. https://pubmed.ncbi.nlm.nih.gov/29667299/
5. Zhang, Xinna, Berger, Franklin G, Yang, Jianhua, Lu, Xiongbin. 2011. USP4 inhibits p53 through deubiquitinating and stabilizing ARF-BP1. In The EMBO journal, 30, 2177-89. doi:10.1038/emboj.2011.125. https://pubmed.ncbi.nlm.nih.gov/21522127/
6. Chen, Cong-Jie, Huang, Jie-Yun, Huang, Jian-Qing, Chen, Long-Tian, Wu, Wei-Hao. 2023. Metformin attenuates multiple myeloma cell proliferation and encourages apoptosis by suppressing METTL3-mediated m6A methylation of THRAP3, RBM25, and USP4. In Cell cycle (Georgetown, Tex.), 22, 986-1004. doi:10.1080/15384101.2023.2170521. https://pubmed.ncbi.nlm.nih.gov/36762777/
7. Xie, Zhongdong, Lin, Hanbin, Wu, Yuecheng, Huang, Ying, Chi, Pan. 2025. USP4-mediated CENPF deubiquitylation regulated tumor metastasis in colorectal cancer. In Cell death & disease, 16, 81. doi:10.1038/s41419-025-07424-3. https://pubmed.ncbi.nlm.nih.gov/39922805/
8. Vlasschaert, Caitlyn, Xia, Xuhua, Coulombe, Josée, Gray, Douglas A. 2015. Evolution of the highly networked deubiquitinating enzymes USP4, USP15, and USP11. In BMC evolutionary biology, 15, 230. doi:10.1186/s12862-015-0511-1. https://pubmed.ncbi.nlm.nih.gov/26503449/
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