MGI:1338027Homozygotes for a null allele die neonatally showing altered patterning of the axial skeleton including abnormal trunk-to-tail transition and ectopic ventral mass with impaired renal, palate, stomach, spleen and pancreatic development. A second null allele alters retinal and olfactory epithelium neurogenesis. A third null allele causes extra thoracic vertebrae.